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The Madagascar animals fly back to New York City, but crash-land on an African nature reserve in Kenya, where they meet others of their own kind, and Alex especially discovers his royal heritage as prince of a lion pride.
The Madagascar animals join a struggling European circus to get back to New York, but find themselves being pursued by a psychotic animal-control officer.
Skipper, Kowalski, Rico and Private join forces with undercover organization The North Wind to stop the villainous Dr. Octavius Brine from destroying the world as we know it.